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  2. Listed below are some of the primary changes in apple physiology that will occur as fruit become more mature:

    • Starch is converted to sugar
    • Seeds turn a darker color
    • Acidity decreases
    • Chlorophyll levels decrease
    • Respiration increases
    • Cell walls weaken, fruit begins to soften
    • Ethylene production increases
    treefruit.wsu.edu/web-article/harvest-apples/
    treefruit.wsu.edu/web-article/harvest-apples/
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